What is the meaning of the word "hackle"?

Definitions:

  1. A tool used for combing flax or hemp fibers to separate the long, desirable fibers from the shorter ones. copy to clipboard NOUN
    • The artisan used a hackle to prepare the flax fibers for spinning. copy to clipboard
  2. Can be used metaphorically to describe a person's state of anger or agitation, often in the phrase 'raise someone's hackles'. copy to clipboard NOUN
    • His rude comments raised her hackles, and she responded sharply. copy to clipboard
  3. The erectile hairs along the back of a dog, cat, or other animal that rise when the animal is agitated or alarmed. copy to clipboard NOUN
    • The dog's hackles rose as it sensed the stranger approaching. copy to clipboard
  4. The neck feathers of a bird, particularly a domestic rooster, which can be raised when the bird is alarmed or displaying aggression. copy to clipboard NOUN
    • The rooster's hackles stood up as it prepared to defend his territory. copy to clipboard

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