What is the meaning of the word "knuckle"?

Definitions:

  1. a joint of a finger when the fist is closed copy to clipboard noun
    • The fighter's knuckles hurt after the cage match. copy to clipboard
  2. to press or rub with the knuckles copy to clipboard verb
    • Can you knuckle my shoulders? copy to clipboard
  3. to shoot a marble while keeping one's knuckles on the ground copy to clipboard verb
    • I knuckle marbles. copy to clipboard

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