What is the meaning of the word "strain"?

Definitions:

  1. (biology) a group of organisms within a species that differ in trivial ways from similar groups copy to clipboard noun
    • a new strain of microorganisms copy to clipboard
  2. difficulty that causes worry or emotional tension copy to clipboard noun
    • I endured the stresses and strains of life. copy to clipboard
  3. nervousness resulting from mental stress copy to clipboard noun
    • The responsibilities were a constant strain. copy to clipboard
  4. to exert much effort or energy copy to clipboard verb
    • straining our ears to hear copy to clipboard

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