What is the meaning of the word "tread"?

Definitions:

  1. crush as if by treading on copy to clipboard verb
    • tread grapes to make wine copy to clipboard
  2. mate with copy to clipboard verb
    • male birds tread the females copy to clipboard
  3. put down or press the foot, place the foot copy to clipboard verb
    • For fools rush in where angels fear to tread copy to clipboard

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